It strongly depends on what functionalities of the JRE are you using, but I'd expect some exceptions, and even strange beaviour in, for example, focus management
It depends entirely on what your applet is doing, and the classes that your code uses.
With minor versions, you will normally be OK, (ie compile on 1.4.2, run on 1.4.1).
With major versions (ie compile on 1.4.2, run on 1.3) - if your code only uses classes that have not changed their class & methods signatures between version, then you will probably be OK, but its uncertain territorary.
Basically, try it and see what happens !
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