In order to filter one box based on the value of the other, the two must share a common field. In essence, you are creating a relationship between the two combo boxes much as you would in a query. Otherwise, what are you filtering on?
In the "RowSource", the fields will display in a manner similar to that of a query. You can pull down any additional fields you wish, such as the common field.
On Current:
When a form is placed in Design view, double-click the gray box that intersects the two rulers (upper left corner). This displays the Properties tab. On the Event tab, click the "On Current" event.
As a side note, the Event tab displays the events that are appropriate for the "selected" object...whether the object is a drop-down box, field, or the form itself.