What DCs you can run depends on the domain and forst functional levels. When you build a new forest and 2008 is your first domain controller the levels are set at:
Domain functional level - windows 2000 native
Forest functional level -windows 2000
Using these levels you can have Windows 2000, 2003 and 2008 domain controllers in the mix.
If you had 2003 and 2008 domain controllers you could then raise these levels to Windows 2003. Here is a link desribing the features of each level;
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