The mailboxes. Commonly /var/spool/mail/*, but maybe somewhers else like the directories under ~user/Maildir. Check the sendmail configuration; the spool files are the usual default location.
In general, assuming that I have both the space and the time, I'd rather just back up everything. Who knows what I might need or want? Better to backup junk I'll never need than miss something.
We use bacula for tape storage, but it also can do stuff to files (local or remote). Bacula is very powerful and useful tool, but it's also a PITA sometimes.
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