Have you specified the path to your information store?
If not look on the server properties in Exchange administrator.Go to "Database Paths" tab and modify to point to the correct database path.
I had a similar problem and it turned ot to be Mcafee groupshield.The engine version of the groupshield was out of date (and for some reason wasnt updating automatically) and was incompatible with the latest DAT version from Mcafee.I manually updated the engine and the problem was gone.
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